Licenciatura / B.Sc.
Universidad Veracruzana, Xalapa, Ver. — Biología-Ecología Terrestre / Biology-Terrestrial Ecology
Doctorado / Ph.D.
University of Missouri, Columbia MO — Ecología, Evolución y Comportamiento / Ecology, Evolution, and Behavior
Posdoctorado / Postdoctoral
Cornell University, Lab of Ornithology — Ithaca NY
Dartmouth College, Biology Department — Hanover NH